/* ==========================================================================
   HOMININES — DESIGN SYSTEM SCIENTIFIQUE, ÉDITORIAL & INTERACTIF (style.css)
   Supporte le Mode Sombre (Observatoire) et le Mode Clair (Édition Parchemin)
   ========================================================================== */

:root {
    /* Palette Mode Sombre (Par défaut : Observatoire Paléoanthropologique) */
    --bg-primary: #090b10;
    --bg-secondary: #10141f;
    --bg-tertiary: #161b2a;
    --bg-card: rgba(20, 25, 38, 0.78);
    --bg-card-solid: #141926;
    --bg-glass: rgba(15, 19, 30, 0.82);
    --bg-glass-border: rgba(255, 255, 255, 0.09);
    --bg-glass-hover: rgba(255, 255, 255, 0.05);

    --text-main: #f1f3f9;
    --text-secondary: #cbd2e1;
    --text-muted: #8e97b0;

    /* Accents scientifiques */
    --accent-cyan: #00d2eb;
    --accent-amber: #f59e0b;
    --accent-terracotta: #e05d44;
    --accent-emerald: #10b981;
    --accent-violet: #a855f7;
    --accent-rose: #ec4899;
    --accent-gold: #eab308;

    /* Couleurs thématiques par grade évolutif (HSL) */
    --h-early: 35;          /* Mio-Pliocène : Ocre / Ambre ancien */
    --h-australo: 145;      /* Australopithèques : Vert savane arborée */
    --h-paranthropus: 25;   /* Paranthropes : Terre cuite robuste */
    --h-habilis: 5;         /* Premiers Homo : Brique Oldowayen */
    --h-erectus: 38;        /* Erectus / Ergaster : Or / Braise */
    --h-antecessor: 162;    /* Antecessor / Casablanca / Heidelbergensis : Turquoise */
    --h-neanderthal: 18;    /* Néandertal : Ocre rouge */
    --h-denisovan: 272;     /* Dénisoviens / Longi : Violet profond */
    --h-sapiens: 190;       /* Homo sapiens : Cyan lumineux */
    --h-island: 320;        /* Insulaires & Naledi : Magenta */

    /* Typographie */
    --font-heading: 'Outfit', -apple-system, BlinkMacSystemFont, sans-serif;
    --font-body: 'Plus Jakarta Sans', -apple-system, BlinkMacSystemFont, sans-serif;
    --font-serif: 'Merriweather', Georgia, serif;

    /* Ombres & Rayons */
    --shadow-card: 0 12px 36px -8px rgba(0, 0, 0, 0.45);
    --shadow-glow: 0 0 28px rgba(0, 210, 235, 0.2);
    --radius-sm: 10px;
    --radius-md: 16px;
    --radius-lg: 24px;
    --radius-xl: 32px;
}

/* Mode Clair ("Édition Scientifique & Collège de France") */
[data-theme="light"] {
    --bg-primary: #f7f4ed;
    --bg-secondary: #efeae0;
    --bg-tertiary: #e5dfd2;
    --bg-card: rgba(255, 253, 249, 0.9);
    --bg-card-solid: #fffdf9;
    --bg-glass: rgba(247, 244, 237, 0.88);
    --bg-glass-border: rgba(40, 35, 25, 0.12);
    --bg-glass-hover: rgba(40, 35, 25, 0.05);

    --text-main: #1b1d24;
    --text-secondary: #333745;
    --text-muted: #5e6478;

    --accent-cyan: #0284c7;
    --accent-amber: #d97706;
    --accent-terracotta: #c2410c;
    --accent-emerald: #059669;
    --accent-violet: #7c3aed;
    --accent-rose: #db2777;
    --accent-gold: #b45309;

    --shadow-card: 0 10px 30px -10px rgba(30, 25, 15, 0.12);
    --shadow-glow: 0 4px 20px rgba(2, 132, 199, 0.15);
}

/* ==========================================================================
   RESET, ACCESSIBILITÉ & BASE
   ========================================================================== */

*, *::before, *::after {
    box-sizing: border-box;
    margin: 0;
    padding: 0;
}

html {
    scroll-behavior: smooth;
    font-size: 16px;
    scroll-padding-top: 90px;
}

body {
    background-color: var(--bg-primary);
    color: var(--text-main);
    font-family: var(--font-body);
    line-height: 1.7;
    overflow-x: hidden;
    position: relative;
    transition: background-color 0.35s ease, color 0.35s ease;
}

.container {
    max-width: 1280px;
    margin: 0 auto;
    padding: 0 24px;
}

a {
    color: inherit;
    text-decoration: none;
}

img {
    max-width: 100%;
    display: block;
}

button, select, input {
    font-family: inherit;
}

/* Barre de progression de lecture */
.reading-progress-bar {
    position: fixed;
    top: 0;
    left: 0;
    height: 3px;
    width: 0%;
    background: linear-gradient(90deg, var(--accent-amber), var(--accent-cyan), var(--accent-violet));
    z-index: 2000;
    transition: width 0.1s linear;
}

/* ==========================================================================
   NAVIGATION STICKY & CONTRÔLES DE THÈME
   ========================================================================== */

.site-nav {
    position: fixed;
    top: 0;
    left: 0;
    right: 0;
    z-index: 1000;
    background: var(--bg-glass);
    backdrop-filter: blur(16px);
    -webkit-backdrop-filter: blur(16px);
    border-bottom: 1px solid var(--bg-glass-border);
    transition: all 0.3s ease;
}

.nav-inner {
    display: flex;
    align-items: center;
    justify-content: space-between;
    height: 74px;
    gap: 16px;
}

.brand-logo {
    display: flex;
    align-items: center;
    gap: 12px;
    font-family: var(--font-heading);
    font-weight: 800;
    font-size: 1.35rem;
    letter-spacing: 0.5px;
}

.brand-icon {
    width: 38px;
    height: 38px;
    border-radius: 10px;
    background: linear-gradient(135deg, var(--accent-amber), var(--accent-cyan));
    display: flex;
    align-items: center;
    justify-content: center;
    font-size: 1.2rem;
    color: #090b10;
    font-weight: 800;
}

.brand-subtitle {
    display: block;
    font-size: 0.7rem;
    font-weight: 500;
    color: var(--text-muted);
    letter-spacing: 1px;
    text-transform: uppercase;
}

.nav-links {
    display: none;
    list-style: none;
    align-items: center;
    gap: 6px;
}

@media (min-width: 1100px) {
    .nav-links {
        display: flex;
    }
}

.nav-links a {
    padding: 8px 13px;
    border-radius: 8px;
    font-size: 0.88rem;
    font-weight: 600;
    color: var(--text-secondary);
    transition: all 0.2s ease;
    white-space: nowrap;
}

.nav-links a:hover,
.nav-links a.active {
    background: var(--bg-glass-hover);
    color: var(--accent-cyan);
}

.nav-actions {
    display: flex;
    align-items: center;
    gap: 10px;
}

.theme-toggle-btn {
    display: inline-flex;
    align-items: center;
    gap: 8px;
    padding: 8px 14px;
    border-radius: 100px;
    background: var(--bg-glass-hover);
    border: 1px solid var(--bg-glass-border);
    color: var(--text-main);
    font-size: 0.85rem;
    font-weight: 600;
    cursor: pointer;
    transition: all 0.2s ease;
}

.theme-toggle-btn:hover {
    border-color: var(--accent-cyan);
    transform: translateY(-1px);
}

/* ==========================================================================
   GLOW BLOBS DÉCORATIFS
   ========================================================================== */

.glow-blob {
    position: fixed;
    border-radius: 50%;
    filter: blur(140px);
    opacity: 0.12;
    z-index: -1;
    pointer-events: none;
}

[data-theme="light"] .glow-blob {
    opacity: 0.07;
}

.glow-1 {
    top: -10%;
    left: -10%;
    width: 45vw;
    height: 45vw;
    background: radial-gradient(circle, var(--accent-cyan), transparent 70%);
}

.glow-2 {
    top: 40%;
    right: -15%;
    width: 45vw;
    height: 45vw;
    background: radial-gradient(circle, var(--accent-amber), transparent 70%);
}

.glow-3 {
    bottom: -10%;
    left: 20%;
    width: 40vw;
    height: 40vw;
    background: radial-gradient(circle, var(--accent-violet), transparent 70%);
}

/* ==========================================================================
   HERO SECTION ÉDITORIALE
   ========================================================================== */

.hero {
    min-height: 92vh;
    padding: 130px 0 80px;
    display: flex;
    align-items: center;
    position: relative;
    overflow: hidden;
}

.hero-bg-banner {
    position: absolute;
    inset: 0;
    background-image: url('assets/header.png');
    background-size: cover;
    background-position: center;
    opacity: 0.16;
    z-index: -1;
    mask-image: linear-gradient(to bottom, rgba(0,0,0,0.8) 30%, transparent 100%);
    -webkit-mask-image: linear-gradient(to bottom, rgba(0,0,0,0.8) 30%, transparent 100%);
}

[data-theme="light"] .hero-bg-banner {
    opacity: 0.22;
}

.hero-grid {
    display: grid;
    grid-template-columns: 1fr;
    gap: 48px;
    align-items: center;
}

@media (min-width: 992px) {
    .hero-grid {
        grid-template-columns: 1.35fr 1fr;
    }
}

.hero-badge {
    display: inline-flex;
    align-items: center;
    gap: 8px;
    background: var(--bg-card);
    border: 1px solid var(--bg-glass-border);
    padding: 8px 18px;
    border-radius: 100px;
    font-size: 0.82rem;
    font-weight: 700;
    letter-spacing: 1.2px;
    text-transform: uppercase;
    color: var(--accent-cyan);
    margin-bottom: 22px;
}

.hero-title {
    font-family: var(--font-heading);
    font-weight: 800;
    font-size: clamp(2.6rem, 5.2vw, 4.4rem);
    line-height: 1.08;
    margin-bottom: 24px;
    letter-spacing: -0.02em;
}

.hero-title .gradient-text {
    background: linear-gradient(135deg, var(--accent-amber) 0%, var(--accent-cyan) 55%, var(--accent-violet) 100%);
    -webkit-background-clip: text;
    background-clip: text;
    -webkit-text-fill-color: transparent;
}

.hero-subtitle {
    font-size: clamp(1.08rem, 1.6vw, 1.28rem);
    color: var(--text-secondary);
    margin-bottom: 28px;
    max-width: 720px;
}

.hero-quote-box {
    background: var(--bg-card);
    border-left: 4px solid var(--accent-amber);
    border-top: 1px solid var(--bg-glass-border);
    border-right: 1px solid var(--bg-glass-border);
    border-bottom: 1px solid var(--bg-glass-border);
    padding: 18px 22px;
    border-radius: 0 var(--radius-md) var(--radius-md) 0;
    margin-bottom: 32px;
}

.hero-quote-text {
    font-family: var(--font-serif);
    font-style: italic;
    font-size: 0.98rem;
    color: var(--text-main);
    margin-bottom: 8px;
}

.hero-quote-author {
    font-size: 0.85rem;
    color: var(--accent-amber);
    font-weight: 700;
}

.hero-actions {
    display: flex;
    flex-wrap: wrap;
    gap: 14px;
}

.btn {
    display: inline-flex;
    align-items: center;
    justify-content: center;
    gap: 8px;
    padding: 13px 24px;
    border-radius: 12px;
    font-family: var(--font-heading);
    font-weight: 700;
    font-size: 0.96rem;
    cursor: pointer;
    transition: all 0.25s ease;
    border: 1px solid transparent;
}

.btn-primary {
    background: linear-gradient(135deg, var(--accent-cyan), #0284c7);
    color: #fff;
    box-shadow: 0 6px 20px rgba(0, 210, 235, 0.25);
}

.btn-primary:hover {
    transform: translateY(-2px);
    box-shadow: 0 10px 26px rgba(0, 210, 235, 0.38);
}

.btn-secondary {
    background: var(--bg-card);
    border-color: var(--bg-glass-border);
    color: var(--text-main);
}

.btn-secondary:hover {
    border-color: var(--accent-cyan);
    transform: translateY(-2px);
}

/* Carte de synthèse chiffrée dans le Hero */
.hero-stats-panel {
    background: var(--bg-card);
    backdrop-filter: blur(18px);
    border: 1px solid var(--bg-glass-border);
    border-radius: var(--radius-lg);
    padding: 28px;
    box-shadow: var(--shadow-card);
    display: flex;
    flex-direction: column;
    gap: 20px;
}

.hero-stats-title {
    font-family: var(--font-heading);
    font-size: 1.05rem;
    font-weight: 700;
    color: var(--accent-cyan);
    text-transform: uppercase;
    letter-spacing: 1px;
    border-bottom: 1px solid var(--bg-glass-border);
    padding-bottom: 12px;
}

.hero-kpi-grid {
    display: grid;
    grid-template-columns: 1fr 1fr;
    gap: 16px;
}

.hero-kpi-card {
    background: var(--bg-glass-hover);
    border: 1px solid var(--bg-glass-border);
    border-radius: var(--radius-md);
    padding: 16px;
}

.hero-kpi-num {
    font-family: var(--font-heading);
    font-size: 1.75rem;
    font-weight: 800;
    color: var(--text-main);
    line-height: 1.1;
}

.hero-kpi-label {
    font-size: 0.8rem;
    color: var(--text-muted);
    margin-top: 6px;
    display: block;
}

/* ==========================================================================
   SECTIONS GÉNÉRIQUES (avec optimisation content-visibility)
   ========================================================================== */

.section {
    padding: 96px 0;
    border-top: 1px solid var(--bg-glass-border);
}

.section-header {
    max-width: 840px;
    margin: 0 auto 56px;
    text-align: center;
}

.section-eyebrow {
    display: inline-block;
    font-family: var(--font-heading);
    font-size: 0.82rem;
    font-weight: 700;
    text-transform: uppercase;
    letter-spacing: 1.6px;
    color: var(--accent-amber);
    margin-bottom: 12px;
}

.section-title {
    font-family: var(--font-heading);
    font-size: clamp(2rem, 3.8vw, 2.9rem);
    font-weight: 800;
    line-height: 1.15;
    margin-bottom: 18px;
}

.section-desc {
    color: var(--text-secondary);
    font-size: 1.08rem;
}

/* ==========================================================================
   SECTION 1 : PROLOGUE — ALAIN PROCHIANTZ ("SINGE TOI-MÊME") & BIFURCATION
   ========================================================================== */

.prochiantz-layout {
    display: grid;
    grid-template-columns: 1fr;
    gap: 32px;
    margin-bottom: 48px;
}

@media (min-width: 992px) {
    .prochiantz-layout {
        grid-template-columns: 1.1fr 0.9fr;
    }
}

.editorial-card {
    background: var(--bg-card);
    border: 1px solid var(--bg-glass-border);
    border-radius: var(--radius-lg);
    padding: 34px;
    box-shadow: var(--shadow-card);
}

.editorial-card h3 {
    font-family: var(--font-heading);
    font-size: 1.55rem;
    font-weight: 800;
    margin-bottom: 16px;
    color: var(--accent-amber);
}

.editorial-card p {
    color: var(--text-secondary);
    margin-bottom: 16px;
    font-size: 1.01rem;
}

.concept-pillars {
    display: grid;
    grid-template-columns: 1fr;
    gap: 16px;
    margin-top: 24px;
}

@media (min-width: 640px) {
    .concept-pillars {
        grid-template-columns: 1fr 1fr;
    }
}

.pillar-box {
    background: var(--bg-glass-hover);
    border: 1px solid var(--bg-glass-border);
    border-radius: var(--radius-md);
    padding: 18px;
}

.pillar-box h4 {
    font-family: var(--font-heading);
    font-size: 1.02rem;
    font-weight: 700;
    color: var(--accent-cyan);
    margin-bottom: 6px;
    display: flex;
    align-items: center;
    gap: 8px;
}

.pillar-box p {
    font-size: 0.9rem;
    color: var(--text-muted);
    margin: 0;
}

/* Schéma interactif Néoténie & Taxonomie */
.interactive-schema-card {
    background: var(--bg-card);
    border: 1px solid var(--bg-glass-border);
    border-radius: var(--radius-lg);
    padding: 28px;
    box-shadow: var(--shadow-card);
    display: flex;
    flex-direction: column;
    justify-content: space-between;
}

.schema-tabs {
    display: flex;
    gap: 8px;
    margin-bottom: 20px;
    flex-wrap: wrap;
}

.schema-tab-btn {
    padding: 8px 16px;
    border-radius: 100px;
    border: 1px solid var(--bg-glass-border);
    background: var(--bg-glass-hover);
    color: var(--text-secondary);
    font-size: 0.85rem;
    font-weight: 700;
    cursor: pointer;
    transition: all 0.2s ease;
}

.schema-tab-btn.active {
    background: var(--accent-cyan);
    color: #090b10;
    border-color: var(--accent-cyan);
}

.schema-pane {
    display: none;
}

.schema-pane.active {
    display: block;
    animation: fadeIn 0.3s ease;
}

.svg-chart-wrapper {
    width: 100%;
    background: var(--bg-secondary);
    border: 1px solid var(--bg-glass-border);
    border-radius: var(--radius-md);
    padding: 16px;
    overflow-x: auto;
}

/* Grille de comparaison Pan vs Homo */
.pan-homo-table {
    width: 100%;
    border-collapse: collapse;
    font-size: 0.9rem;
    margin-top: 12px;
}

.pan-homo-table th,
.pan-homo-table td {
    padding: 10px 12px;
    text-align: left;
    border-bottom: 1px solid var(--bg-glass-border);
}

.pan-homo-table th {
    color: var(--accent-amber);
    font-family: var(--font-heading);
    font-weight: 700;
}

/* ==========================================================================
   SECTION 2 : ARBRE PHYLOGÉNÉTIQUE BUISSONNANT INTERACTIF (SVG)
   ========================================================================== */

.phylo-controls-bar {
    display: flex;
    flex-wrap: wrap;
    align-items: center;
    justify-content: space-between;
    gap: 16px;
    margin-bottom: 24px;
    background: var(--bg-card);
    border: 1px solid var(--bg-glass-border);
    padding: 16px 24px;
    border-radius: var(--radius-md);
}

.phylo-legend {
    display: flex;
    flex-wrap: wrap;
    gap: 14px;
    align-items: center;
}

.legend-item {
    display: inline-flex;
    align-items: center;
    gap: 7px;
    font-size: 0.84rem;
    color: var(--text-secondary);
    font-weight: 600;
}

.legend-dot {
    width: 12px;
    height: 12px;
    border-radius: 50%;
    display: inline-block;
}

.phylo-stage-wrapper {
    background: var(--bg-card);
    border: 1px solid var(--bg-glass-border);
    border-radius: var(--radius-lg);
    padding: 24px;
    box-shadow: var(--shadow-card);
    overflow-x: auto;
}

#phylo-svg-container {
    min-width: 920px;
    width: 100%;
}

.phylo-node-group {
    cursor: pointer;
    transition: transform 0.25s ease, opacity 0.25s ease;
}

.phylo-node-group:hover circle.node-main-circle {
    r: 22px;
    filter: drop-shadow(0 0 12px currentColor);
}

.phylo-link {
    fill: none;
    stroke-width: 2.5px;
    opacity: 0.65;
    transition: opacity 0.25s ease, stroke-width 0.25s ease;
}

.phylo-hybrid-link {
    fill: none;
    stroke-width: 2.2px;
    stroke-dasharray: 6 5;
    animation: flowDash 20s linear infinite;
}

@keyframes flowDash {
    to {
        stroke-dashoffset: -220;
    }
}

/* ==========================================================================
   SECTION 3 : MACHINE À REMONTER LE TEMPS & COURBE D'ENCÉPHALISATION
   ========================================================================== */

.timemachine-panel {
    background: var(--bg-card);
    border: 1px solid var(--bg-glass-border);
    border-radius: var(--radius-lg);
    padding: 32px;
    box-shadow: var(--shadow-card);
    margin-bottom: 48px;
}

.preset-epochs-bar {
    display: flex;
    flex-wrap: wrap;
    gap: 10px;
    margin-bottom: 24px;
    justify-content: center;
}

.epoch-preset-btn {
    padding: 8px 14px;
    border-radius: 100px;
    border: 1px solid var(--bg-glass-border);
    background: var(--bg-glass-hover);
    color: var(--text-secondary);
    font-size: 0.83rem;
    font-weight: 600;
    cursor: pointer;
    transition: all 0.2s ease;
}

.epoch-preset-btn:hover,
.epoch-preset-btn.active {
    background: var(--accent-amber);
    color: #090b10;
    border-color: var(--accent-amber);
}

.time-slider-row {
    display: flex;
    align-items: center;
    gap: 18px;
    margin-bottom: 20px;
}

#global-time-slider {
    flex: 1;
    -webkit-appearance: none;
    appearance: none;
    height: 8px;
    border-radius: 4px;
    background: linear-gradient(90deg, var(--accent-amber), var(--accent-emerald), var(--accent-cyan));
    outline: none;
}

#global-time-slider::-webkit-slider-thumb {
    -webkit-appearance: none;
    appearance: none;
    width: 24px;
    height: 24px;
    border-radius: 50%;
    background: #fff;
    border: 4px solid var(--accent-cyan);
    cursor: pointer;
    box-shadow: 0 0 12px rgba(0, 210, 235, 0.7);
}

.time-readout-banner {
    display: flex;
    flex-wrap: wrap;
    align-items: center;
    justify-content: space-between;
    gap: 16px;
    padding: 18px 24px;
    background: var(--bg-secondary);
    border-radius: var(--radius-md);
    border: 1px solid var(--bg-glass-border);
    margin-bottom: 24px;
}

.time-readout-main {
    font-family: var(--font-heading);
    font-size: 1.5rem;
    font-weight: 800;
    color: var(--accent-cyan);
}

.time-coexistence-badge {
    padding: 6px 14px;
    border-radius: 100px;
    background: rgba(16, 185, 129, 0.15);
    border: 1px solid rgba(16, 185, 129, 0.4);
    color: var(--accent-emerald);
    font-weight: 700;
    font-size: 0.9rem;
}

.active-species-strip {
    display: grid;
    grid-template-columns: repeat(auto-fill, minmax(240px, 1fr));
    gap: 16px;
}

.mini-species-pill {
    background: var(--bg-glass-hover);
    border: 1px solid var(--bg-glass-border);
    border-radius: var(--radius-md);
    padding: 14px 16px;
    display: flex;
    align-items: center;
    gap: 12px;
    cursor: pointer;
    transition: all 0.2s ease;
}

.mini-species-pill:hover {
    border-color: var(--accent-cyan);
    transform: translateY(-2px);
}

.mini-species-avatar {
    width: 46px;
    height: 46px;
    border-radius: 50%;
    object-fit: cover;
    flex-shrink: 0;
    background: var(--bg-secondary);
    border: 2px solid var(--accent-cyan);
    display: flex;
    align-items: center;
    justify-content: center;
    font-weight: 800;
    font-size: 0.85rem;
}

/* ==========================================================================
   SECTION 4 : FOCUS JEAN-JACQUES HUBLIN (773 ka CASABLANCA → JEBEL IRHOUD)
   ========================================================================== */

.hublin-spotlight {
    background: linear-gradient(145deg, var(--bg-card), var(--bg-secondary));
    border: 1px solid var(--bg-glass-border);
    border-radius: var(--radius-xl);
    padding: 40px;
    box-shadow: var(--shadow-card);
}

.hublin-milestones-grid {
    display: grid;
    grid-template-columns: 1fr;
    gap: 24px;
    margin: 32px 0;
}

@media (min-width: 900px) {
    .hublin-milestones-grid {
        grid-template-columns: repeat(3, 1fr);
    }
}

.milestone-card {
    background: var(--bg-card);
    border: 1px solid var(--bg-glass-border);
    border-top: 4px solid var(--accent-cyan);
    border-radius: var(--radius-md);
    padding: 26px;
    position: relative;
}

.milestone-card:nth-child(1) {
    border-top-color: var(--accent-amber);
}

.milestone-card:nth-child(2) {
    border-top-color: var(--accent-emerald);
}

.milestone-date {
    font-family: var(--font-heading);
    font-size: 1.65rem;
    font-weight: 800;
    color: var(--text-main);
    margin-bottom: 6px;
}

.milestone-site {
    display: inline-block;
    font-size: 0.82rem;
    font-weight: 700;
    text-transform: uppercase;
    letter-spacing: 0.8px;
    color: var(--accent-cyan);
    margin-bottom: 14px;
}

.milestone-card p {
    font-size: 0.94rem;
    color: var(--text-secondary);
}

/* ==========================================================================
   SECTION 5 : ENCYCLOPÉDIE DES 16 ESPÈCES (FILTRABLE)
   ========================================================================== */

.filter-toolbar {
    display: flex;
    flex-wrap: wrap;
    gap: 12px;
    justify-content: space-between;
    align-items: center;
    margin-bottom: 36px;
}

.filter-pills {
    display: flex;
    flex-wrap: wrap;
    gap: 8px;
}

.filter-btn {
    padding: 9px 18px;
    border-radius: 100px;
    border: 1px solid var(--bg-glass-border);
    background: var(--bg-card);
    color: var(--text-secondary);
    font-family: var(--font-heading);
    font-size: 0.88rem;
    font-weight: 600;
    cursor: pointer;
    transition: all 0.2s ease;
}

.filter-btn:hover,
.filter-btn.active {
    background: var(--accent-cyan);
    color: #090b10;
    border-color: var(--accent-cyan);
}

.search-input-box {
    padding: 10px 18px;
    border-radius: 100px;
    border: 1px solid var(--bg-glass-border);
    background: var(--bg-card);
    color: var(--text-main);
    font-size: 0.92rem;
    min-width: 260px;
    outline: none;
}

.search-input-box:focus {
    border-color: var(--accent-cyan);
}

.species-encyclopedia-grid {
    display: grid;
    grid-template-columns: 1fr;
    gap: 28px;
}

@media (min-width: 768px) {
    .species-encyclopedia-grid {
        grid-template-columns: repeat(2, 1fr);
    }
}

@media (min-width: 1140px) {
    .species-encyclopedia-grid {
        grid-template-columns: repeat(3, 1fr);
    }
}

.species-card {
    background: var(--bg-card);
    border: 1px solid var(--bg-glass-border);
    border-radius: var(--radius-lg);
    overflow: hidden;
    display: flex;
    flex-direction: column;
    box-shadow: var(--shadow-card);
    transition: transform 0.3s ease, border-color 0.3s ease;
}

.species-card:hover {
    transform: translateY(-6px);
    border-color: var(--accent-cyan);
}

.species-visual {
    position: relative;
    height: 230px;
    background: var(--bg-secondary);
    overflow: hidden;
    display: flex;
    align-items: center;
    justify-content: center;
}

.species-visual img {
    width: 100%;
    height: 100%;
    object-fit: cover;
    transition: transform 0.5s ease;
}

.species-card:hover .species-visual img {
    transform: scale(1.06);
}

.species-svg-placeholder {
    width: 100%;
    height: 100%;
    display: flex;
    flex-direction: column;
    align-items: center;
    justify-content: center;
    background: radial-gradient(circle at center, rgba(0, 210, 235, 0.14), transparent 75%);
    padding: 20px;
    text-align: center;
}

.species-visual-badges {
    position: absolute;
    bottom: 12px;
    left: 12px;
    right: 12px;
    display: flex;
    justify-content: space-between;
    align-items: center;
    gap: 8px;
}

.badge-pill {
    padding: 5px 11px;
    border-radius: 100px;
    font-size: 0.76rem;
    font-weight: 700;
    background: rgba(9, 11, 16, 0.82);
    color: #fff;
    border: 1px solid rgba(255, 255, 255, 0.15);
    backdrop-filter: blur(6px);
}

.species-card-body {
    padding: 24px;
    display: flex;
    flex-direction: column;
    flex: 1;
}

.species-card-title {
    font-family: var(--font-heading);
    font-size: 1.4rem;
    font-weight: 800;
    font-style: italic;
    margin-bottom: 4px;
}

.species-card-sub {
    font-size: 0.85rem;
    color: var(--accent-amber);
    font-weight: 600;
    margin-bottom: 14px;
}

.species-card-desc {
    font-size: 0.93rem;
    color: var(--text-secondary);
    margin-bottom: 18px;
    flex: 1;
}

.species-metrics-row {
    display: grid;
    grid-template-columns: repeat(3, 1fr);
    gap: 8px;
    padding: 12px 0;
    border-top: 1px solid var(--bg-glass-border);
    border-bottom: 1px solid var(--bg-glass-border);
    margin-bottom: 16px;
}

.metric-cell {
    display: flex;
    flex-direction: column;
}

.metric-val {
    font-family: var(--font-heading);
    font-weight: 700;
    font-size: 0.95rem;
    color: var(--text-main);
}

.metric-lbl {
    font-size: 0.7rem;
    text-transform: uppercase;
    color: var(--text-muted);
}

.species-card-actions {
    display: flex;
    justify-content: space-between;
    align-items: center;
    gap: 10px;
}

.btn-sm {
    padding: 8px 14px;
    font-size: 0.83rem;
    border-radius: 8px;
}

/* ==========================================================================
   SECTION 6 : COMPARATEUR MULTI-ESPÈCES & PALÉOGÉNOMIQUE
   ========================================================================== */

.comparator-wrapper {
    background: var(--bg-card);
    border: 1px solid var(--bg-glass-border);
    border-radius: var(--radius-xl);
    padding: 36px;
    box-shadow: var(--shadow-card);
}

.comparator-controls {
    display: flex;
    flex-wrap: wrap;
    align-items: center;
    justify-content: center;
    gap: 24px;
    margin-bottom: 36px;
}

.control-group {
    display: flex;
    flex-direction: column;
    gap: 8px;
    min-width: 260px;
}

.control-group label {
    font-size: 0.78rem;
    text-transform: uppercase;
    letter-spacing: 1.2px;
    color: var(--text-muted);
    font-weight: 700;
}

.select-species {
    background-color: var(--bg-secondary);
    border: 1px solid var(--bg-glass-border);
    color: var(--text-main);
    padding: 12px 16px;
    border-radius: 12px;
    font-family: var(--font-heading);
    font-weight: 700;
    font-size: 0.98rem;
    cursor: pointer;
}

.vs-divider {
    font-family: var(--font-heading);
    font-weight: 800;
    font-size: 1.3rem;
    color: var(--accent-amber);
    padding: 10px 16px;
    border-radius: 50%;
    background: var(--bg-glass-hover);
    border: 1px solid var(--bg-glass-border);
}

.comparison-grid {
    display: grid;
    grid-template-columns: 1fr;
    gap: 28px;
}

@media (min-width: 820px) {
    .comparison-grid {
        grid-template-columns: 1fr 1fr;
    }
}

.compare-column {
    background: var(--bg-secondary);
    border: 1px solid var(--bg-glass-border);
    padding: 28px;
    border-radius: var(--radius-lg);
}

.compare-card-head {
    display: flex;
    align-items: center;
    gap: 18px;
    margin-bottom: 24px;
    padding-bottom: 18px;
    border-bottom: 1px solid var(--bg-glass-border);
}

.compare-card-head .img-wrapper {
    width: 76px;
    height: 76px;
    border-radius: 50%;
    overflow: hidden;
    border: 2px solid var(--accent-cyan);
    flex-shrink: 0;
    background: var(--bg-primary);
    display: flex;
    align-items: center;
    justify-content: center;
}

.compare-card-head img {
    width: 100%;
    height: 100%;
    object-fit: cover;
}

.compare-stats-list {
    display: flex;
    flex-direction: column;
    gap: 16px;
}

.compare-stat-row {
    display: flex;
    flex-direction: column;
    gap: 4px;
}

.compare-lbl {
    font-size: 0.76rem;
    text-transform: uppercase;
    letter-spacing: 0.9px;
    color: var(--text-muted);
}

.compare-val {
    font-family: var(--font-heading);
    font-weight: 700;
    font-size: 1.08rem;
    color: var(--text-main);
}

.brain-bar-container {
    width: 100%;
    height: 9px;
    background: rgba(128, 128, 128, 0.18);
    border-radius: 5px;
    overflow: hidden;
    margin-top: 4px;
}

.brain-bar {
    height: 100%;
    width: 0%;
    border-radius: 5px;
    transition: width 0.6s cubic-bezier(0.25, 0.8, 0.25, 1);
}

/* ==========================================================================
   SECTION 7 : PALÉOANTHROPOLOGUES, GÉNÉTICIENS & INSTITUTIONS
   ========================================================================== */

.scholars-grid,
.institutions-grid,
.videos-grid,
.glossary-grid {
    display: grid;
    grid-template-columns: 1fr;
    gap: 24px;
}

@media (min-width: 768px) {
    .scholars-grid,
    .institutions-grid,
    .videos-grid,
    .glossary-grid {
        grid-template-columns: repeat(2, 1fr);
    }
}

@media (min-width: 1120px) {
    .scholars-grid,
    .institutions-grid,
    .videos-grid,
    .glossary-grid {
        grid-template-columns: repeat(3, 1fr);
    }
}

.scholar-card,
.institution-card,
.video-card,
.glossary-card {
    background: var(--bg-card);
    border: 1px solid var(--bg-glass-border);
    border-radius: var(--radius-md);
    padding: 24px;
    box-shadow: var(--shadow-card);
    display: flex;
    flex-direction: column;
    justify-content: space-between;
    transition: transform 0.25s ease, border-color 0.25s ease;
}

.scholar-card:hover,
.institution-card:hover,
.video-card:hover,
.glossary-card:hover {
    transform: translateY(-4px);
    border-color: var(--accent-cyan);
}

.scholar-role-badge {
    display: inline-block;
    font-size: 0.74rem;
    font-weight: 700;
    text-transform: uppercase;
    letter-spacing: 0.8px;
    padding: 4px 10px;
    border-radius: 100px;
    background: var(--bg-glass-hover);
    color: var(--accent-cyan);
    border: 1px solid var(--bg-glass-border);
    margin-bottom: 12px;
    align-self: flex-start;
}

.scholar-name {
    font-family: var(--font-heading);
    font-size: 1.3rem;
    font-weight: 800;
    margin-bottom: 4px;
}

.scholar-affiliation {
    font-size: 0.85rem;
    color: var(--accent-amber);
    font-weight: 600;
    margin-bottom: 12px;
}

.scholar-bio {
    font-size: 0.92rem;
    color: var(--text-secondary);
    margin-bottom: 14px;
}

.scholar-works {
    font-size: 0.84rem;
    color: var(--text-muted);
    padding-top: 12px;
    border-top: 1px solid var(--bg-glass-border);
    margin-bottom: 12px;
}

.ext-link {
    display: inline-flex;
    align-items: center;
    gap: 6px;
    font-size: 0.86rem;
    font-weight: 700;
    color: var(--accent-cyan);
}

.ext-link:hover {
    text-decoration: underline;
}

/* ==========================================================================
   SECTION 8 : MÉDIATHÈQUE VIDÉOS YOUTUBE & CONFÉRENCES
   ========================================================================== */

.video-thumb-box {
    position: relative;
    border-radius: var(--radius-sm);
    overflow: hidden;
    background: linear-gradient(135deg, #161f33, #0d1322);
    aspect-ratio: 16 / 9;
    margin-bottom: 16px;
    display: flex;
    flex-direction: column;
    align-items: center;
    justify-content: center;
    padding: 18px;
    text-align: center;
    border: 1px solid var(--bg-glass-border);
    cursor: pointer;
}

.video-thumb-box img {
    position: absolute;
    inset: 0;
    width: 100%;
    height: 100%;
    object-fit: cover;
    opacity: 0.55;
    transition: opacity 0.3s ease, transform 0.4s ease;
}

.video-card:hover .video-thumb-box img {
    opacity: 0.75;
    transform: scale(1.04);
}

.play-circle-btn {
    position: relative;
    z-index: 2;
    width: 56px;
    height: 56px;
    border-radius: 50%;
    background: rgba(224, 93, 68, 0.92);
    color: #fff;
    display: flex;
    align-items: center;
    justify-content: center;
    font-size: 1.35rem;
    box-shadow: 0 6px 20px rgba(0, 0, 0, 0.5);
    transition: transform 0.25s ease;
}

.video-card:hover .play-circle-btn {
    transform: scale(1.12);
}

.video-duration-tag {
    position: absolute;
    bottom: 10px;
    right: 10px;
    z-index: 2;
    background: rgba(0, 0, 0, 0.8);
    color: #fff;
    padding: 3px 8px;
    border-radius: 6px;
    font-size: 0.75rem;
    font-weight: 700;
}

.youtube-privacy-notice {
    margin-top: 28px;
    padding: 14px 20px;
    border-radius: var(--radius-sm);
    background: var(--bg-secondary);
    border: 1px solid var(--bg-glass-border);
    font-size: 0.82rem;
    color: var(--text-muted);
    text-align: center;
}

/* ==========================================================================
   SECTION 9 : QUIZ PÉDAGOGIQUE INTERACTIF
   ========================================================================== */

.quiz-container {
    max-width: 820px;
    margin: 0 auto;
    background: var(--bg-card);
    border: 1px solid var(--bg-glass-border);
    border-radius: var(--radius-lg);
    padding: 36px;
    box-shadow: var(--shadow-card);
}

.quiz-header-bar {
    display: flex;
    justify-content: space-between;
    align-items: center;
    margin-bottom: 20px;
    padding-bottom: 14px;
    border-bottom: 1px solid var(--bg-glass-border);
}

.quiz-question-text {
    font-family: var(--font-heading);
    font-size: 1.35rem;
    font-weight: 700;
    margin-bottom: 22px;
}

.quiz-options-list {
    display: flex;
    flex-direction: column;
    gap: 12px;
    margin-bottom: 22px;
}

.quiz-option-btn {
    text-align: left;
    padding: 14px 18px;
    border-radius: var(--radius-sm);
    border: 1px solid var(--bg-glass-border);
    background: var(--bg-secondary);
    color: var(--text-main);
    font-size: 0.96rem;
    font-weight: 500;
    cursor: pointer;
    transition: all 0.2s ease;
}

.quiz-option-btn:hover:not(:disabled) {
    border-color: var(--accent-cyan);
    background: var(--bg-glass-hover);
}

.quiz-option-btn.correct {
    border-color: var(--accent-emerald);
    background: rgba(16, 185, 129, 0.16);
}

.quiz-option-btn.wrong {
    border-color: var(--accent-terracotta);
    background: rgba(224, 93, 68, 0.16);
}

.quiz-feedback-box {
    padding: 16px 20px;
    border-radius: var(--radius-sm);
    background: var(--bg-secondary);
    border-left: 4px solid var(--accent-cyan);
    font-size: 0.93rem;
    color: var(--text-secondary);
    margin-bottom: 20px;
    display: none;
}

/* ==========================================================================
   MODALE NATIVE <dialog> POUR LES FICHES DÉTAILLÉES & VIDÉOS
   ========================================================================== */

dialog.species-dialog {
    margin: auto;
    width: min(92vw, 940px);
    max-height: 88vh;
    border: 1px solid var(--bg-glass-border);
    border-radius: var(--radius-lg);
    background: var(--bg-card-solid);
    color: var(--text-main);
    box-shadow: 0 25px 60px rgba(0, 0, 0, 0.65);
    padding: 0;
    overflow: hidden;
}

dialog.species-dialog::backdrop {
    background: rgba(5, 7, 12, 0.78);
    backdrop-filter: blur(8px);
}

.dialog-inner {
    display: grid;
    grid-template-columns: 1fr;
    max-height: 88vh;
    overflow-y: auto;
}

@media (min-width: 780px) {
    .dialog-inner.has-image {
        grid-template-columns: 0.9fr 1.1fr;
    }
}

.dialog-visual-col {
    background: var(--bg-secondary);
    min-height: 280px;
    position: relative;
    display: flex;
    align-items: center;
    justify-content: center;
}

.dialog-visual-col img {
    width: 100%;
    height: 100%;
    object-fit: cover;
}

.dialog-content-col {
    padding: 32px;
    overflow-y: auto;
}

.dialog-top-bar {
    display: flex;
    justify-content: space-between;
    align-items: flex-start;
    gap: 16px;
    margin-bottom: 18px;
}

.dialog-close-btn {
    width: 38px;
    height: 38px;
    border-radius: 50%;
    border: 1px solid var(--bg-glass-border);
    background: var(--bg-glass-hover);
    color: var(--text-main);
    font-size: 1.2rem;
    cursor: pointer;
    flex-shrink: 0;
}

.dialog-close-btn:hover {
    border-color: var(--accent-terracotta);
    color: var(--accent-terracotta);
}

/* ==========================================================================
   FOOTER
   ========================================================================== */

.footer {
    border-top: 1px solid var(--bg-glass-border);
    background: var(--bg-secondary);
    padding: 60px 0 40px;
    color: var(--text-muted);
    font-size: 0.92rem;
}

.footer-grid {
    display: grid;
    grid-template-columns: 1fr;
    gap: 32px;
    margin-bottom: 36px;
}

@media (min-width: 768px) {
    .footer-grid {
        grid-template-columns: 1.5fr 1fr 1fr;
    }
}

.footer h4 {
    font-family: var(--font-heading);
    color: var(--text-main);
    margin-bottom: 12px;
    font-size: 1.05rem;
}

.footer ul {
    list-style: none;
    display: flex;
    flex-direction: column;
    gap: 8px;
}

.footer a:hover {
    color: var(--accent-cyan);
}

/* ==========================================================================
   BOUTON FLOTTANT RETOUR EN HAUT
   ========================================================================== */

.back-to-top-btn {
    position: fixed;
    bottom: 28px;
    right: 28px;
    z-index: 1500;
    width: 48px;
    height: 48px;
    border-radius: 50%;
    background: linear-gradient(135deg, var(--accent-cyan), #0284c7);
    color: #fff;
    border: 1px solid rgba(255, 255, 255, 0.25);
    font-family: var(--font-heading);
    font-size: 1.35rem;
    font-weight: 800;
    display: flex;
    align-items: center;
    justify-content: center;
    cursor: pointer;
    box-shadow: 0 8px 24px rgba(0, 210, 235, 0.35);
    opacity: 0;
    visibility: hidden;
    transform: translateY(14px);
    transition: opacity 0.3s ease, transform 0.3s ease, visibility 0.3s ease, box-shadow 0.25s ease;
}

.back-to-top-btn.visible {
    opacity: 1;
    visibility: visible;
    transform: translateY(0);
}

.back-to-top-btn:hover {
    transform: translateY(-3px);
    box-shadow: 0 12px 28px rgba(0, 210, 235, 0.55);
}

@keyframes fadeIn {
    from { opacity: 0; transform: translateY(6px); }
    to { opacity: 1; transform: translateY(0); }
}